1. Plymouth Marine Laboratory, Prospect Place, The Hoe, Plymouth PL1 3DH,
United Kingdom
2. College of Life and Environmental
Sciences: Biosciences, Geoffrey Pope Building, University of Exeter, Stocker Road, Exeter EX4 4QD, United Kingdom
3. Akvaplan-niva AS, FRAM − High North Research Centre for Climate and the Environment, N-9296 Tromsø, Norway
4. College of Engineering, Mathematics
and Physical Sciences: Physics, Physics Building, University of Exeter, Stocker Road, Exeter EX4 4QL, United Kingdom