1. The Laboratory of Medical Genetics, Harbin Medical University, Harbin 150081, China
2. The 2th Affiliated Hospital, Harbin Medical University, Harbin 150001, China
3. Key Laboratory of Preservation of Human Genetic Resources and Disease Control in China (Harbin Medical University), Ministry of Education, Harbin150081, China
4. Department of Cell Biology, Harvard Medical School, Boston, Massachusetts 02115, United States