1. Section of Plant Pathology and Plant-Microbe Biology, School of Integrated Plant Sciences, Cornell University, Ithaca, New York 14853, United States
2. Boyce Thompson Institute, Ithaca, New York 14853, United States
3. Emerging Pests and Pathogens Research Unit, Robert W. Holley Center, USDA ARS, Ithaca, New York 14853, United States