1. Department of Ophthalmology, West Virginia University, Morgantown, West Virginia 26506, United States
2. Department of Biochemistry, West Virginia University, Morgantown, West Virginia 26506, United States
3. Save Sight Institute, Sydney Medical School, University of Sydney, Sydney, NSW 2000, Australia
4. Department of Ophthalmology, University of Washington, Seattle, Washington 98109, United States