1. Department of Biochemistry and Cell Biology, Geisel School of Medicine at Dartmouth, Hanover, New Hampshire NH 03755, United States
2. Department of Molecular and Systems Biology, Geisel School of Medicine at Dartmouth, Hanover, New Hampshire NH 03755, United States
3. Dartmouth Cancer Center, Geisel School of Medicine at Dartmouth, Lebanon, New Hampshire 03756, United States