1. Fisheries College, Guangdong Ocean University, Zhanjiang 524025, China
2. Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy Culture and Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Zhanjiang 524025, Guangdong, China
3. Chinese Academy of Inspection and Quarantine, Beijing 100176, China
4. School of Public Health, Fujian Medical University, Fujian 350122, China