1. South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Guangzhou 510300, China
2. Faculty of Science, Yamagata University, Yamagata 990-8560, Japan
3. Guangdong Provincial Key Laboratory of Fishery Ecology and Environment, Guangzhou 510300, China
4. Center for Ecological Environment Monitoring and Scientific Research, Huaihe Valley Ecology and Environment Administration,Ministry of Ecology and Environment, Bengbu 233000, China
5. College of Oceanography, Hohai University, Nanjing 210024, China