1. Center for Marine Environmental Studies (CMES), Ehime University, Bunkyo-cho 2-5, Matsuyama 790-8577, Japan, Branch Office in Ogasawara, Tokyo Metropolitan Government, Nishimachi, Chichijima, Ogasawara, Tokyo 100-2101, Japan, and National Research Institute for Fishery Environment of Inland Sea, Fishery Research Agency, Maruishi 2-17-5, Ohno-Cho, Hiroshima 739-0452, Japan