1. Instituto de Biomedicina de Valencia, C.S.I.C., Jaume Roig 11, 46010 Valencia, Spain, Department of Evolution, Ecology and Organismal Biology, Ohio State University, 300 Aronoff Laboratory, 318 West 12th Avenue, Columbus, Ohio 43210-1293, and School of Biological Sciences, University of Northern Colorado, 501 20th Street, CB 92, Greeley, Colorado 80639-0017