1. Center for Biophysics and Quantitative Biology, University of Illinois at Urbana−Champaign, Urbana, Illinois 61820, United States
2. Department of Physics, University of Illinois at Urbana−Champaign, Urbana, Illinois 61820, United States
3. Center for the Physics of Living Cells, University of Illinois at Urbana−Champaign, Urbana, Illinois 61820, United States
4. Department of Biophysics, Johns Hopkins University, Baltimore, Maryland 21218, United States
5. Program in Cell, Molecular, Developmental Biology, and Biophysics, Johns Hopkins University, Baltimore, Maryland 21218, United States