1. Department of Bioresource Engineering, Yamagata University, Tsuruoka, Yamagata 997-8555, Japan, Division of Chemistry, Graduate School of Science, Hokkaido University, Sapporo 060-0810, Japan, Biotechnology Research Center, Toyama Prefectural University, Imizu, Toyoma 939-0398, Japan, and The Institute of Scientific and Industrial Research, Osaka University, Ibaraki, Osaka 567-0047, Japan