1. Okazaki Institute for Integrative Bioscience, National Institutes of Natural Sciences, 5-1 Higashiyama, Myodaiji, Okazaki, 444-8787, Japan, University of Exeter, School of Biosciences, Prince of Wales Road, Exeter, Devon, EX4 4PS, United Kingdom, and Brunel University, Department of Biological Sciences, Uxbridge, Middlesex UB8 3PH, United Kingdom