1. College of Biotechnology, Tianjin University of Science and Technology, Tianjin 300457, China
2. Key Laboratory of Engineering Biology for Low-carbon Manufacturing, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, China
3. Key Laboratory of Engineering Biology for Low-carbon Manufacturing, Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, National Center of Technology Innovation for Synthetic Biology, Tianjin 300308, China
4. Tianjin Zhonghe Gene Technology Company Limited, Tianjin 300308, China