1. Department of Chemical and Biomolecular Engineering (BK21 Four), Korea Advanced Institute of Science and Technology (KAIST), Daejeon 34141, Republic of Korea
2. Center for Infectious Disease Vaccine and Diagnosis Innovation, Korea Research Institute of Chemical Technology (KRICT), Daejeon 34114, Republic of Korea
3. Center for Systems Biology, Massachusetts General Hospital, Boston, Massachusetts 02114, United States
4. Department of Radiology, Massachusetts General Hospital, Harvard Medical School, Boston, Massachusetts 02114, United States