1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao 266237, People’s Republic of China
2. Institute of Marine Science and Technology, Shandong University, Qingdao 266237, People’s Republic of China
3. School of Life Science, Shandong University, Qingdao 266237, People’s Republic of China
4. School of Molecular Biosciences, Washington State University, Pullman, Washington 99164-7520, United States