1. Department of Biochemistry and Biophysics, University of Pennsylvania, Philadelphia, Pennsylvania 19104-6243, United States
2. Epigenetic Institute and Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, Pennsylvania 19104-6243, United States
3. Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, St. Louis, Missouri 63110-1010, United States
4. Department of Biochemistry, Albert Einstein College of Medicine, Bronx, New York 10461-1900, United States