1. South China Institute of Botany, The Chinese Academy of Sciences, Guangzhou ReYiJu 510650, People's Republic of China; School of Land and Food Sciences, The University of Queensland, Brisbane, Queensland 4072, Australia; Department of Agro-Industry, Faculty of Agriculture, Natural Resources and Environment, Naresuan University, Muang, Phitsanulok 65000, Thailand; and School of Agriculture and Horticulture, The University of Queensland, Gatton, Queensland 4343, Australia