1. Graduate School of Biomedical Sciences, Hiroshima University, 1-2-3 Kasumi, Minami-ku, Hiroshima 734-8553, Japan, Subtropical Field Science Center, Faculty of Agriculture, University of the Ryukyus, 1 Sembaru, Nishihara-cho, Nakagami-gun, Okinawa 903-0213, Japan, and Faculty of Integrated Arts and Sciences, The University of Tokushima, 1 Minamijosanjima-cho, Tokushima 770-8502, Japan