1. Laboratory of Biological Chemistry, School of Marine Biosciences, Kitasato University, Sagamihara, Kanagawa 252-0373, Japan
2. Laboratory of Aquatic Molecular Biology and Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo-ku, Tokyo 113-8657, Japan
3. Sugashima Marine Biological Laboratory, Graduate School of Science, Nagoya University, Toba, Mie 517-0001, Japan