1. Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Key Laboratory of Marine Materia Medica, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Sanya Institute of Ocean Eco-Environmental Engineering, Yazhou Scientific Bay, Sanya, 572000, China
4. Department of Agricultural Biology, Faculty of Agriculture, University of Ruhuna, Matara 81000, Sri Lanka