1. Graduate School of Bioagricultural Sciences, Nagoya University, Chikusa-ku, Nagoya 464-8601, Japan, Graduate School of Biomedical Sciences, Hiroshima University, Minami-ku, Hiroshima 734-8553, Japan, ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, Queensland 4811, Australia, and National Institute for Environmental Studies, Onogawa, Tsukuba 305-8506, Japan