1. Centre for Human Development, Stem Cells and Regeneration, Institute for Developmental Sciences, Southampton General Hospital, University of Southampton, Tremona Road, Southampton SO16 6YD, Hampshire, U.K.
2. School of Biological Sciences, University of Southampton, Highfield Campus, University Road, Southampton SO17 1BJ, Hampshire, U.K.
3. Faculty of Health Sciences, University of Southampton, Highfield Campus, University Road, Southampton SO17 1BJ, U.K.