1. School of Food Biosciences, The University of Reading, P.O. Box 226, Reading RG6 6AP, United Kingdom; Brewing Research International, Lyttel Hall, Nutfield, Surrey RH1 4HY, United Kingdom; and Dipartimento di Scienze degli Alimenti, Università di Udine, via Marangoni 97, I-33100 Udine, Italy