1. Guelph Waterloo Program for Graduate Studies in Physics and Department of Biology, University of Waterloo, Waterloo, Ontario N2L 3G1, Canada, Radiation Oncology, E-1200, Vanderbilt Medical Centre North, Nashville, Tennessee 37232, and Radiation Oncology/Physics/Research Laboratory, William Beaumont Hospital, 3601 West 13 Mile Road, Royal Oak, Michigan 48073