1. Ehime Prefectural Institute of Public Health and Environmental Science, Sanbancho 8-234, Matsuyama, Ehime 790-0003, Japan, Center for Marine Environmental Studies (CMES), Ehime University, Bunkyo-cho 2-5, Matsuyama 790-8577, Japan, and Otsuchi Marine Research Center, Ocean Research Institute, The University of Tokyo, Akahama, Otsuchi-cho, Iwate 028-1102, Japan