1. Brookdale Department of Molecular, Cell, and Developmental Biology, Mount Sinai School of Medicine, One Gustave L. Levy Place, New York, New York 10029-6574, and Institut fuer Molekulare Medizin und Zellforschung, Albert-Ludwigs Universitaet, Stefan-Meier-Strasse 19, 79104 Freiburg, Germany