1. Biotechnology Research Center, Toyama Prefectural University, 5180 Kurokawa, Imizu, Toyama 939-0398, Japan, National Institute of Infectious Diseases, 1-23-1 Toyama, Shinjuku, Tokyo 162-8640, Japan, Department of Bioscience, Institute of Natural Medicine,University of Toyama, 2630 Sugitani, Toyama 930-0194, Japan, and Departmento de Microbiologia y Genetica, Edificio Departamental Lab. 205, Campus Miguel de Unamuno, Universidad de Salamanca, 37007 Salamanca, Spain