1. Bristol Dental School, University of Bristol, Bristol BS1 2LY, U.K.
2. National Physical Laboratory, Teddington TW11 0LW, U.K.
3. Department of Biology, College of Science, University of Jeddah, Jeddah 23218, Saudi Arabia
4. Centre for the Cellular Microenvironment, University of Glasgow, Glasgow G11 6EW, Scotland