1. Program in Cell, Molecular, Developmental Biology, and Biophysics, Johns Hopkins University, Baltimore, Maryland 21218, United States
2. Department of Biology, Johns Hopkins University, Baltimore, Maryland 21218, United States
3. Department of Biochemistry and Biophysics, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania 19104, United States
4. Department of Biophysics, Johns Hopkins University, Baltimore, Maryland 21218, United States
5. Physics Frontier Center (Center for the Physics of Living Cells), University of Illinois at Urbana−Champaign, Urbana, Illinois 61801, United States