1. MRC Center for Drug Safety Science, Department of Molecular and Clinical Pharmacology, University of Liverpool, Sherrington Building, Ashton Street, Liverpool, L69 3GE, England
2. Institute of Cellular Medicine, Medical School, Newcastle University, Newcastle NE2 4HH, United Kingdom
3. GlaxoSmithKline, Park Road, Ware, Hertfordshire SG12 7BB, United Kingdom