1. Colorado State
University, Department of Fish, Wildlife, and Conservation Biology, Fort Collins, Colorado 80523 United States
2. University of South Dakota, Department of Biology, Vermillion, South Dakota 57069 United States
3. U.S. Geological Survey, Crustal Geophysics and
Geochemistry Science Center, Denver, Colorado 80225, United States
4. U.S. Geological
Survey, Fort Collins Science Center, 2150 Centre Ave - Bldg C, Fort Collins, Colorado 80525 United States