1. Institute for Molecular Bioscience, The University of Queensland, Brisbane 4072, Queensland, Australia
2. Whitehead Institute for Biomedical Research, 9 Cambridge Center, Cambridge, Massachusetts 02142, United States
3. Program in Cellular and Molecular Medicine, Division of Molecular Biology, Department of Medicine, Boston Children’s Hospital, 3 Blackfan Circle, Third Floor, Boston, Massachusetts 02115, United States