1. Department of Bioengineering, Graduate School of Engineering, The University of Tokyo, Tokyo 113-8656, Japan
2. Institute of Medical Science, The University of Tokyo, Tokyo 108-8639, Japan
3. Laboratory of Global Healthcare, Graduate School of Pharmaceutical Sciences, Kyushu University, Fukuoka 812-8582, Japan
4. Department of Material and Biological Chemistry, Faculty of Science, Yamagata University, Yamagata 990-8560, Japan
5. Ubiquitin Project, Tokyo Metropolitan Institute of Medical Science, 2-1-6 Kamikitazawa, Setagaya, Tokyo 156-8506, Japan
6. Department of Chemistry and Biochemistry, Loyola University Chicago, Chicago, Illinois 60660, United States
7. Department of Chemistry and Biotechnology, School of Engineering, The University of Tokyo, Tokyo 108-8639, Japan