1. Department of Biological Chemistry and Molecular Pharmacology, Blavatnik Institute, Harvard Medical School, Boston Massachusetts 02115, United States
2. Broad Institute of MIT and Harvard, Cambridge, Massachusetts 02142, United States
3. Department of Biological Sciences, Virginia Tech, Blacksburg, Virginia 24061, United States
4. Department of Molecular Biology, Massachusetts General Hospital, Boston, Massachusetts 02114, United States
5. Center for the Study of Inflammatory Bowel Disease, Massachusetts General Hospital, Boston, Massachusetts 02114, United States