1. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo 113-8657, Japan
2. Collaborative Research Institute for Innovative Microbiology (CRIIM), The University of Tokyo, 1-1-1 Yayoi, Bunkyo, Tokyo 113-8657, Japan
3. New Field Pioneering Division, Toyota Boshoku Corporation, 1-1 Toyoda, Kariya, Aichi 448-8651, Japan
4. Graduate School of Medicine, Nagoya University, 65 Tsurumai, Showa, Nagoya 466-8550, Japan