1. Department of Cell Biology, Johns Hopkins University, Baltimore, Maryland 21205 United States
2. Johns Hopkins Drug Discovery and Department of Neurology, Johns Hopkins University, Baltimore, Maryland 21205 United States
3. Department of Oncology, Sidney Kimmel Comprehensive Cancer Center, Johns Hopkins University,Baltimore, Maryland 21205 United States
4. Giovanis Institute for Translational Cell Biology, Johns Hopkins University School of Medicine, Baltimore, Maryland 21205 United States