1. University of Exeter, Biosciences, College of Life & Environmental Sciences, Exeter EX4 4PS, United Kingdom
2. Okazaki Institute for Integrative Bioscience, National Institute for Basic Biology, National Institutes of Natural Sciences, and Department of Basic Biology, School of Life Science, Graduate University for Advanced Studies, 5-1 Higashiyama, Myodaiji, Okazaki 444-8787, Japan