1. Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, St. Louis, Missouri 63110, United States
2. Center for Science and Engineering Living Systems (CSELS), Washington University, St. Louis, Missouri 63130, United States
3. Centre for Genomic Regulation (CRG), The Barcelona Institute of Science and Technology, Dr. Aiguader 88, Barcelona 08003, Spain
4. Universitat Pompeu Fabra (UPF), Barcelona 08002, Spain