1. Graduate School of Life Sciences, Ritsumeikan University, 1-1-1 Noji-higashi, Kusatsu, Shiga 525-8577, Japan
2. College of Life Sciences, Ritsumeikan University, 1-1-1 Noji-higashi, Kusatsu, Shiga 525-8577, Japan
3. Graduate School of Information Science, University of Hyogo, 7-1-28 minatojima Minamimachi, Chuo-ku, Kobe, Hyogo 650-0047, Japan
4. Research Organization of Science and Technology, Ritsumeikan University, 1-1-1 Noji-higashi, Kusatsu, Shiga 525-8577, Japan