1. School of Biomedical Sciences, The University of Queensland, St Lucia, Brisbane, QLD 4072, Australia
2. University of Queensland Centre for Clinical Research, the University of Queensland, Brisbane, QLD 4029, Australia
3. Queensland Brain Institute, the University of Queensland, St Lucia, Brisbane, QLD 4072, Australia
4. Wesley Medical Research, The Wesley Hospital, Auchenflower, Brisbane, QLD 4066, Australia