1. Research Faculty of Agriculture, Division of Fundamental Agriscience Research, Hokkaido University, North-9, West-9, Kita-ku, Sapporo, Hokkaido 060-8589, Japan
2. Faculty of Biological Science and Technology, Institute of Science and Engineering, Kanazawa University, Kakuma-machi, Kanazawa, Ishikawa 920-1192, Japan