1. Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology, Qingdao 266237, China
3. National and Local Joint Engineering Laboratory of Ecological Mariculture, Qingdao 266071, China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
5. Laboratory for Marine Fisheries and Aquaculture, Pilot National Laboratory for Marine Science and Technology, Qingdao 266237, China
6. The Innovation of Seed Design, Chinese Academy of Sciences, Wuhan 430072, P. R. China