1. School of Life Sciences, Liaocheng University, Liaocheng, Shandong 252000, China
2. State Key Laboratory of Marine Resource Utilization in the South China Sea, School of Marine Biology and Aquaculture, Hainan University, Haikou, Hainan 570100, China
3. Xiamen Key Laboratory of Urban Sea Ecological Conservation and Restoration, State Key Laboratory of Marine Environmental Science, College of Ocean and Earth Sciences, Xiamen University, Xiamen, Fujian 361005, China
4. Key Laboratory of Engineering Biology for Low-Carbon Manufacturing, Tianjin Institute of Industrial Biotech-nology, Chinese Academy of Sciences, Tianjin 300308, China