1. College of Plant Protection, Hunan Agricultural University, Changsha 410128, China
2. Institute of Agricultural Environment and Ecology, Hunan academy of Agricultural Sciences, Changsha 410125, China
3. Department of Crop and Soil Sciences, North Carolina State University, Raleigh, North Carolina 27695, United States
4. Key Laboratory for Biology and Control of Weeds, Hunan Agricultural Biotechnology Research Institute, Hunan Academy of Agricultural Sciences, Changsha 410125, China