1. College of Forest Sciences, Kangwon National University, Chunchon 200-701, Korea, Graduate School of Science, Kyoto University, Kitashirakawa Oiwakecho, Kyoto 606-8502, Japan, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Tokyo 113-8657, Japan, and Research Institute for Sustainable Humanosphere, Kyoto University, Uji, Kyoto 611-0011, Japan