1. Key Laboratory of Marine Environmental Science and Ecology, Ministry of Education, Ocean University of China, Qingdao 266100, China
2. Qingdao National Laboratory for Marine Science and Technology, Qingdao 266003, China
3. Centre for Ocean and Climate Research, First Institute of Oceanography, State Oceanic Administration, Qingdao 266061, China
4. School of Marine Science and Policy, University of Delaware, Newark, Delaware 19716, United States