1. Key Laboratory of Ministry of Education for Coastal Wetland Ecosystems, College of the Environment and Ecology, Xiamen University, Xiamen, Fujian 361102, China
2. Xiamen Key Laboratory of Urban Sea Ecological Conservation and Restoration (USER), Xiamen University, Xiamen, Fujian 361102, China
3. Fujian Provincial Key Laboratory for Coastal Ecology and Environmental Studies, Xiamen University, Xiamen, Fujian 361102, China
4. Coastal and Ocean Management Institute, Xiamen University, Xiamen, Fujian 361102, China
5. Department of Biological & Marine Sciences, University of Hull, Hull HU6 7RX, United Kingdom
6. Third Institute of Oceanography, Ministry of Natural Resources, 178 Daxue Road, Siming District, Xiamen, Fujian 361005, China