1. U.S. Geological Survey, Western Ecological Research Center, Dixon Field Station, Dixon, California 95620, United States of America
2. Institute of Marine Sciences, University of California Santa Cruz, Santa Cruz, California 95064, United States of America
3. Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, California 95064, United States of America
4. Moss Landing Marine Labs, San Jose State University, Moss Landing, California 95039, United States of America
5. Department of Biology, Sonoma State University, Rohnert Park, California 94928, United States of America