1. CAS Key Laboratory of Biofuels, Shandong Provincial Key Laboratory of Energy Genetics, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao 266101, China
2. Shandong Energy Institute, Qingdao 266101, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. State Key Laboratory of Heavy Oil Processing, China University of Petroleum, Beijing 102249, China
5. Qingdao Municipal Bureau of Ecology and Environment, Qingdao 266003, China
6. Institute of Marine Microbes and Ecospheres, State Key Laboratory of Marine Environmental Science, Xiamen University, Xiamen 361100, China