1. Eco&Sols, INRAE, IRD, Cirad, Institut Agro, Univ Montpellier, 34090 Montpellier, France
2. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization and Ecological Restoration Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China
3. SPO, Institut Agro, INRAE, Univ Montpellier, 34090 Montpellier, France